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Beta Diversity Calculator

Calculate beta diversity and community similarity

Beta Diversity Inputs

The Beta Diversity Calculator helps quantify the difference in species composition between two different sites or communities. It uses common ecological indices like Jaccard and Sørensen to measure similarity or dissimilarity.

Beta diversity is a key component of biodiversity, reflecting how species composition changes across a landscape.

Jaccard Index = (Number of shared species) / (Total number of species in both sites)

Sørensen Index = (2 * Number of shared species) / (Total species in Site A + Total species in Site B)

Enter species lists to calculate beta diversity

About Beta Diversity Calculator

Unveiling Ecological Differences: The Power of Beta Diversity

In the vast tapestry of life, biodiversity is often discussed in terms of species richness within a single area (alpha diversity) or the total number of species across a large region (gamma diversity). However, a crucial dimension of biodiversity, often overlooked but equally vital for understanding ecological patterns and processes, is beta diversity. Beta diversity quantifies the difference in species composition between two or more sites or communities. It tells us not just how many species are present, but how those species change from one location to another, providing insights into habitat heterogeneity, environmental gradients, and the spatial dynamics of ecological communities.

Our Beta Diversity Calculator is a powerful tool designed to help ecologists, conservationists, and students quantify these differences using widely accepted ecological indices. By inputting species lists from two distinct sites, you can quickly calculate metrics like the Jaccard Index and Sørensen Index, gaining a clearer picture of community similarity or dissimilarity. This understanding is fundamental for effective conservation planning, ecological monitoring, and research into the drivers of biodiversity patterns.

What is Beta Diversity and Why Does it Matter?

Imagine two forests. Both might have the same number of tree species (similar alpha diversity). But if one forest has oak, maple, and pine, and the other has birch, spruce, and fir, their species compositions are entirely different. Beta diversity captures this difference. It is a measure of species turnover or compositional change along environmental gradients or between different habitats.

The importance of beta diversity extends across various ecological disciplines:

  • Conservation Planning: Identifying areas with high beta diversity can highlight regions that contribute uniquely to overall biodiversity, making them priorities for protection. It also helps in designing protected area networks that capture a wide range of habitats and species assemblages.
  • Environmental Monitoring: Changes in beta diversity over time can indicate environmental degradation, habitat fragmentation, or the impact of climate change on species distributions.
  • Understanding Ecological Processes: Beta diversity patterns can reveal the influence of dispersal limitation, environmental filtering, and species interactions in shaping community structure.
  • Biogeography: It helps explain why certain species are found in some regions but not others, contributing to our understanding of large-scale biodiversity patterns.
  • Restoration Ecology: Assessing beta diversity before and after restoration efforts can gauge the success of interventions in re-establishing characteristic species assemblages.

Common Measures of Beta Diversity: Jaccard and Sørensen Indices

There are numerous indices to quantify beta diversity, each with its own nuances and applications. Our calculator focuses on two of the most widely used and intuitive similarity indices:

Jaccard Index (Jaccard Similarity Coefficient)

The Jaccard Index measures the similarity between two sets of data. In ecology, it quantifies the proportion of shared species relative to the total number of species found in both sites combined. It ranges from 0 (no shared species, completely dissimilar) to 1 (all species are shared, identical communities).

Jaccard Index (J) = |A ∩ B| / |A ∪ B|
Where: A = species in Site A, B = species in Site B, |A ∩ B| = number of shared species, |A ∪ B| = total number of unique species in both sites.

Sørensen Index (Sørensen-Dice Coefficient)

The Sørensen Index is another popular similarity measure that gives more weight to shared species. It is calculated as twice the number of shared species divided by the sum of the number of species in each site. Like Jaccard, it ranges from 0 (no shared species) to 1 (identical communities).

Sørensen Index (S) = 2 * |A ∩ B| / (|A| + |B|)
Where: A = species in Site A, B = species in Site B, |A ∩ B| = number of shared species, |A| = number of species in Site A, |B| = number of species in Site B.

While both indices measure similarity, the Sørensen Index is often preferred in ecological studies because it is less sensitive to sample size differences and gives more weight to species that are common to both sites. However, the choice of index often depends on the specific research question and the nature of the data.

How Our Beta Diversity Calculator Works

Our calculator makes the computation of these indices straightforward. Here's how to use it:

  • Input Species for Site A: Enter the list of species observed in your first study site. Species names should be separated by commas (e.g., "Oak, Maple, Pine, Birch"). The order does not matter, and duplicate entries within a single site will be treated as unique.
  • Input Species for Site B: Similarly, enter the list of species observed in your second study site, also separated by commas (e.g., "Pine, Spruce, Fir, Maple").

Once you've entered both lists, click 'Calculate Beta Diversity'. The tool will then:

  • Identify the unique species present in each site.
  • Determine the number of species shared between both sites (intersection).
  • Calculate the total number of unique species across both sites (union).
  • Compute the Jaccard Index and Sørensen Index based on these values.

The results will be displayed numerically and visually through a bar chart, allowing for quick comparison of the similarity indices. This visual representation helps in understanding the degree of overlap and uniqueness between the two ecological communities.

Interpreting Your Beta Diversity Results

The values obtained from the Jaccard and Sørensen indices provide a quantitative measure of community similarity:

  • Values close to 1: Indicate high similarity between the two sites. This means the communities share a large proportion of their species, suggesting similar environmental conditions, good connectivity, or a broad species distribution.
  • Values close to 0: Indicate low similarity (high dissimilarity) between the two sites. This suggests that the communities have very few or no species in common, possibly due to different environmental conditions, geographical barriers, or distinct ecological histories.

For example, if the Jaccard Index is 0.2, it means only 20% of the total species found across both sites are shared. If it's 0.8, then 80% are shared. These values can be compared across different pairs of sites or over time for the same sites to track ecological changes.

Factors Influencing Beta Diversity

Beta diversity patterns are shaped by a complex interplay of ecological and evolutionary processes:

Environmental Heterogeneity

Variations in environmental conditions (e.g., soil type, moisture, temperature, elevation) across a landscape lead to different species being favored in different areas, thus increasing beta diversity.

Geographic Distance and Dispersal Limitation

As distance between sites increases, beta diversity often increases because species dispersal becomes limited, preventing them from colonizing all suitable habitats. Barriers like mountains or rivers can also limit dispersal.

Habitat Fragmentation

The breaking up of continuous habitats into smaller, isolated patches can increase beta diversity by creating distinct, isolated communities, but it can also lead to overall biodiversity loss.

Species Interactions

Competition, predation, and mutualism can influence which species coexist in a given area, contributing to differences in community composition between sites.

Historical Factors

Past events like glaciation, geological changes, or historical land use can leave a legacy that influences current species distributions and beta diversity patterns.

Disturbance Regimes

The frequency, intensity, and type of disturbances (e.g., fire, flood, logging) can create mosaics of habitats at different successional stages, thereby influencing beta diversity.

Applications of Beta Diversity in Ecology and Conservation

The insights gained from beta diversity analysis have practical applications across various fields:

  • Reserve Design: Conservation planners use beta diversity to ensure that protected area networks capture a representative sample of regional biodiversity, not just areas with high alpha diversity.
  • Impact Assessment: Evaluating changes in beta diversity before and after human disturbances (e.g., urbanization, deforestation) can quantify the ecological impact of such activities.
  • Invasive Species Management: High beta diversity can sometimes indicate resistance to invasive species, while a decrease might signal homogenization of communities due to invaders.
  • Climate Change Research: Monitoring shifts in beta diversity can help track how species distributions are changing in response to climate change, indicating range contractions or expansions.
  • Restoration Success: Comparing the beta diversity of restored sites with reference sites can assess how well restoration efforts have re-established natural community patterns.
  • Biomonitoring: Using beta diversity as an indicator of ecosystem health, where significant deviations from expected patterns might signal environmental stress.

Beyond Jaccard and Sørensen: Other Beta Diversity Concepts

While our calculator focuses on presence-absence based similarity indices, it's worth noting that beta diversity can also be measured using abundance data (e.g., Bray-Curtis dissimilarity) or phylogenetic relationships (phylogenetic beta diversity), which consider the evolutionary relatedness of species. There are also different conceptualizations of beta diversity, such as:

  • Turnover: The replacement of one species by another across sites.
  • Nestedness: When species in less diverse sites are a subset of species in more diverse sites.
  • Gradient Analysis: Examining how species composition changes along environmental gradients (e.g., elevation, moisture).

These advanced concepts provide a more nuanced understanding of the processes driving biodiversity patterns, but the Jaccard and Sørensen indices remain excellent starting points for fundamental comparisons.

Conclusion: A Window into Ecological Connectivity and Change

The Beta Diversity Calculator offers a straightforward yet powerful means to explore the spatial dimension of biodiversity. By quantifying the differences in species composition between communities, it provides invaluable insights into the factors shaping ecological patterns, from local habitat variations to broad biogeographical processes.

Understanding beta diversity is not merely an academic exercise; it is a critical component of effective environmental stewardship. As habitats continue to transform under human influence and climate change, tools like this calculator empower researchers, conservationists, and land managers to make more informed decisions, ensuring that our efforts to protect biodiversity are as comprehensive and impactful as possible. We encourage you to experiment with different species lists, explore the resulting similarity values, and deepen your appreciation for the intricate and dynamic nature of ecological communities.

Frequently Asked Questions

What is beta diversity?
Beta diversity quantifies the difference in species composition between two or more sites or communities. It measures species turnover or compositional change along environmental gradients or between different habitats.
Why is beta diversity important in ecology?
Beta diversity is crucial because it helps understand how species composition changes across a landscape, revealing patterns of habitat heterogeneity, environmental gradients, and the spatial dynamics of ecological communities. It's vital for conservation planning and monitoring.
How does beta diversity differ from alpha and gamma diversity?
Alpha diversity is the species richness within a single site or community. Gamma diversity is the total species richness across a large region or landscape. Beta diversity links these two, measuring the differentiation between habitats or communities within a region.
What is the Jaccard Index?
The Jaccard Index (Jaccard Similarity Coefficient) is a common measure of similarity between two sets of data. In ecology, it quantifies the proportion of shared species relative to the total number of unique species found in both sites combined.
How is the Jaccard Index calculated?
Jaccard Index = (Number of shared species) / (Total number of unique species in both sites). It ranges from 0 (no shared species) to 1 (identical communities).
What is the Sørensen Index?
The Sørensen Index (Sørensen-Dice Coefficient) is another popular similarity measure that gives more weight to shared species. It is calculated as twice the number of shared species divided by the sum of the number of species in each site.
How is the Sørensen Index calculated?
Sørensen Index = (2 * Number of shared species) / (Total species in Site A + Total species in Site B). Like Jaccard, it ranges from 0 to 1.
Which index, Jaccard or Sørensen, is generally preferred in ecological studies and why?
The Sørensen Index is often preferred in ecological studies because it is less sensitive to sample size differences and gives more weight to species that are common to both sites, making it robust for presence-absence data.
What does a Jaccard or Sørensen index value close to 1 indicate?
A value close to 1 indicates high similarity between the two sites, meaning the communities share a large proportion of their species. This suggests similar environmental conditions or good connectivity.
What does a Jaccard or Sørensen index value close to 0 indicate?
A value close to 0 indicates low similarity (high dissimilarity) between the two sites, suggesting that the communities have very few or no species in common, possibly due to different environmental conditions or barriers.
How do I input species data into the calculator?
You should input species names for each site, separated by commas (e.g., 'Oak, Maple, Pine'). The order does not matter, and the calculator will automatically identify unique species.
Can I use common names or scientific names for species input?
You can use either common names or scientific names, but it's crucial to be consistent within your dataset to ensure accurate matching of shared species. For example, always use 'Oak' or always use 'Quercus'.
What if there are duplicate species entries within a single site's input?
The calculator treats duplicate entries within a single site as unique species for the purpose of counting. It's best practice to list each unique species only once per site.
How does environmental heterogeneity influence beta diversity?
Environmental heterogeneity (variations in conditions like soil, moisture, temperature) across a landscape leads to different species being favored in different areas, thereby increasing beta diversity.
How does geographic distance affect beta diversity?
As the geographic distance between sites increases, beta diversity often increases because species dispersal becomes limited, preventing them from colonizing all suitable habitats and leading to more distinct communities.
What is the role of dispersal limitation in shaping beta diversity?
Dispersal limitation occurs when species cannot reach all suitable habitats due to barriers or insufficient dispersal capabilities. This restricts their distribution and contributes to differences in species composition between sites, thus increasing beta diversity.
How does habitat fragmentation impact beta diversity?
Habitat fragmentation can increase beta diversity by creating distinct, isolated communities in the remaining patches. However, it can also lead to an overall loss of regional biodiversity if species are unable to persist in smaller, isolated fragments.
Can species interactions influence beta diversity?
Yes, competition, predation, and mutualism can influence which species can coexist in a given area. These interactions can lead to different community compositions in different sites, contributing to beta diversity.
What are some applications of beta diversity in conservation planning?
Beta diversity helps in designing protected area networks that capture a representative sample of regional biodiversity, identifying unique habitats, and prioritizing areas for conservation that contribute most to overall landscape diversity.
How can beta diversity be used in environmental monitoring?
Changes in beta diversity over time can indicate environmental degradation, habitat fragmentation, or the impact of climate change on species distributions, making it a valuable metric for long-term environmental monitoring programs.
What is species turnover?
Species turnover is a component of beta diversity that refers to the replacement of one species by another across sites or along an environmental gradient. It highlights the spatial change in species composition.
What is nestedness in the context of beta diversity?
Nestedness occurs when species in less diverse sites are a subset of species found in more diverse sites. It suggests a hierarchical pattern of species loss or gain, often seen in fragmented landscapes.
How can beta diversity analysis inform restoration ecology?
By comparing the beta diversity of restored sites with reference (undisturbed) sites, restoration ecologists can assess the success of their efforts in re-establishing characteristic species assemblages and ecological patterns.
What is phylogenetic beta diversity?
Phylogenetic beta diversity considers the evolutionary relatedness of species when comparing communities. It measures how much evolutionary history is shared or unshared between sites, providing a deeper insight into community assembly.
How does climate change affect beta diversity?
Climate change can alter environmental conditions, leading to species range shifts, extinctions, and colonizations. These processes can change species composition across landscapes, thereby impacting beta diversity patterns.
What is the relationship between beta diversity and ecosystem function?
High beta diversity can contribute to ecosystem stability and function by ensuring that a wider range of species and functional traits are present across a landscape, providing redundancy and resilience to disturbances.
Can beta diversity be used to identify biodiversity hotspots?
While alpha diversity (species richness) is often used to define hotspots, areas with high beta diversity are also crucial as they represent unique species assemblages and contribute disproportionately to regional biodiversity.
What are some limitations of using only Jaccard or Sørensen indices?
These indices are based on presence-absence data and do not account for species abundances. They also treat all species equally, regardless of their ecological role or evolutionary history.
How can beta diversity be measured using abundance data?
Indices like the Bray-Curtis dissimilarity index are commonly used to measure beta diversity when species abundance data is available. These indices account for both species presence and their relative abundances.
What is the concept of 'spatial turnover' in beta diversity?
Spatial turnover refers to the extent to which species composition changes as one moves across a landscape. It's a direct measure of how different communities are from each other in space.
How do environmental gradients influence beta diversity patterns?
Environmental gradients (e.g., elevation, moisture, nutrient availability) create varying conditions that filter species, leading to distinct species assemblages along the gradient and thus high beta diversity.
What is the role of historical factors in shaping beta diversity?
Past geological events, climate shifts, or human land-use history can leave a lasting legacy on species distributions and community assembly, influencing current beta diversity patterns.
How does disturbance regime affect beta diversity?
The frequency, intensity, and type of disturbances (e.g., fire, flood) can create mosaics of habitats at different successional stages, promoting heterogeneity in species composition and thus influencing beta diversity.
What is the relationship between beta diversity and ecosystem resilience?
High beta diversity can contribute to ecosystem resilience by ensuring that a diverse set of species and functional groups are distributed across the landscape, allowing the ecosystem to better withstand and recover from disturbances.
How can beta diversity be used in biomonitoring?
Beta diversity can serve as an indicator of ecosystem health. Significant deviations from expected beta diversity patterns (e.g., homogenization) might signal environmental stress or degradation.
What is the concept of 'community homogenization'?
Community homogenization is a process where the beta diversity of a region decreases, leading to communities becoming more similar to each other over time, often due to human activities like habitat destruction, species introductions, or pollution.
How does species pool size affect beta diversity?
A larger regional species pool (gamma diversity) generally provides more potential species to draw from, which can lead to higher beta diversity if environmental conditions allow for differentiation among sites.
What is the importance of scale in beta diversity studies?
The scale at which beta diversity is measured (e.g., local, regional, continental) significantly influences the observed patterns and the ecological processes that are most relevant. Different scales reveal different insights.
How does habitat connectivity influence beta diversity?
Increased habitat connectivity (e.g., through corridors) can facilitate species dispersal, potentially leading to a decrease in beta diversity as communities become more similar due to increased gene flow and colonization.
What is the relationship between beta diversity and ecosystem services?
Beta diversity can underpin the provision of ecosystem services by ensuring a diverse array of species and functional groups are present across a landscape, contributing to processes like pollination, pest control, and nutrient cycling.
How can beta diversity be used to assess the impact of invasive species?
Invasive species can reduce beta diversity by outcompeting native species and homogenizing communities across different sites. Monitoring beta diversity can help track the ecological impact of invasions.
What is the concept of 'functional beta diversity'?
Functional beta diversity measures the turnover of functional traits (e.g., feeding habits, reproductive strategies) between communities, providing insights into how ecosystem functions change across a landscape.
How does land use change affect beta diversity?
Land use change (e.g., urbanization, agriculture) often leads to habitat loss and fragmentation, which can significantly alter beta diversity patterns, typically by increasing dissimilarity between remaining patches or by homogenizing disturbed areas.
What is the role of 'sampling effort' in beta diversity estimation?
Adequate and standardized sampling effort is crucial for accurate beta diversity estimation. Insufficient sampling can lead to an overestimation of dissimilarity due to undetected species.
How does beta diversity contribute to regional biodiversity?
Beta diversity is a direct component of regional (gamma) diversity. Gamma diversity = Alpha diversity * Beta diversity. High beta diversity means that even if local richness is moderate, the overall regional richness can be very high due to species turnover.
What is the concept of 'species sorting'?
Species sorting is a process where environmental conditions filter species from the regional species pool, allowing only those with suitable traits to persist in a given local community. This contributes to beta diversity.
How does 'environmental filtering' relate to beta diversity?
Environmental filtering occurs when abiotic conditions (e.g., temperature, pH) exclude species that lack the necessary physiological tolerances. This process leads to distinct species assemblages in different environments, thus increasing beta diversity.
What is the difference between 'dissimilarity' and 'similarity' indices?
Similarity indices (like Jaccard and Sørensen) measure how alike two communities are, with values closer to 1 indicating more similarity. Dissimilarity indices (e.g., Jaccard dissimilarity = 1 - Jaccard similarity) measure how different they are, with values closer to 1 indicating more dissimilarity.
How can beta diversity be visualized?
Beta diversity can be visualized using various methods, including ordination techniques (e.g., Non-metric Multidimensional Scaling - NMDS, Principal Components Analysis - PCA) which plot communities based on their species composition, or through heatmaps and dendrograms.
What is the importance of 'null models' in beta diversity research?
Null models are used to test whether observed beta diversity patterns are significantly different from what would be expected by random chance. They help distinguish between deterministic processes (e.g., environmental filtering) and stochastic processes (e.g., dispersal limitation).
How does 'succession' influence beta diversity over time?
Ecological succession involves changes in species composition over time following a disturbance. As communities progress through successional stages, beta diversity can change, often peaking at intermediate stages due to a mix of early and late successional species.
What is the concept of 'beta-diversity partitioning'?
Beta-diversity partitioning involves breaking down total beta diversity into additive components, such as turnover (species replacement) and nestedness (species loss/gain), to understand the relative importance of different processes driving community differences.
How does 'sampling bias' affect beta diversity estimates?
Sampling bias, such as unequal sampling effort or incomplete inventories, can lead to inaccurate estimates of beta diversity, often overestimating dissimilarity due to missed species.
What is the role of 'metacommunities' in beta diversity?
A metacommunity is a set of local communities linked by dispersal of multiple interacting species. Metacommunity theory explores how dispersal, environmental filtering, and species interactions jointly determine beta diversity patterns across a landscape.
How does 'trophic structure' influence beta diversity?
Differences in trophic structure (e.g., food web complexity, presence of top predators) between sites can lead to distinct species assemblages and thus contribute to beta diversity, as different trophic interactions shape community composition.
What is the significance of 'rare species' in beta diversity?
Rare species, even if present in only one site, contribute to beta diversity by increasing the dissimilarity between communities. Their presence can highlight unique habitats or the influence of dispersal limitation.
How can beta diversity be used to prioritize areas for ecological restoration?
Prioritizing restoration efforts in areas that contribute significantly to regional beta diversity can help maximize the recovery of overall biodiversity and ensure that unique species assemblages are re-established.
What is the concept of 'biotic homogenization'?
Biotic homogenization is the process by which the biological communities in different places become more similar over time, often due to human activities. It results in a decrease in beta diversity and a loss of unique local biotas.
How does 'habitat specialization' affect beta diversity?
Species that are highly specialized to particular habitats will contribute more to beta diversity, as they are likely to be found only in those specific conditions, leading to greater differentiation between communities.
What is the role of 'species pools' in determining beta diversity?
The regional species pool (the set of all species available to colonize local communities) sets the upper limit for beta diversity. The processes of dispersal and environmental filtering then determine which species from this pool establish in each local community.
How does 'sampling effort' affect beta diversity estimates?
Inadequate or unequal sampling effort can lead to biased estimates of beta diversity. If one site is undersampled, it might appear more dissimilar than it truly is, as many species might be missed.
What is the concept of 'beta-diversity decomposition'?
Beta-diversity decomposition is a method to break down total beta diversity into components related to species replacement (turnover) and species richness differences (nestedness), providing a more detailed understanding of the underlying ecological processes.
How does 'environmental stress' influence beta diversity?
Environmental stress can reduce the number of species that can tolerate certain conditions, leading to a decrease in alpha diversity. However, if different stressors affect different sites, it can also increase beta diversity by creating distinct, stress-adapted communities.
What is the significance of 'endemism' in beta diversity?
Endemic species (found only in a specific area) contribute significantly to beta diversity, as their unique presence makes a community distinct from others. Areas with high endemism are often conservation priorities.
How does 'succession' influence beta diversity over time?
Ecological succession, the process of change in species structure over time, can influence beta diversity. Early and late successional stages often have different species compositions, leading to higher beta diversity across a landscape with varying successional stages.
What is the role of 'habitat corridors' in managing beta diversity?
Habitat corridors facilitate species movement between fragmented patches, which can reduce beta diversity by promoting community similarity. However, they are crucial for maintaining overall biodiversity and genetic flow in fragmented landscapes.
How does 'trophic structure' influence beta diversity?
Differences in trophic structure (e.g., food web complexity, presence of top predators) between sites can lead to distinct species assemblages and thus contribute to beta diversity, as different trophic interactions shape community composition.
What is the significance of 'rare species' in beta diversity?
Rare species, even if present in only one site, contribute to beta diversity by increasing the dissimilarity between communities. Their presence can highlight unique habitats or the influence of dispersal limitation.
How can beta diversity be used to prioritize areas for ecological restoration?
Prioritizing restoration efforts in areas that contribute significantly to regional beta diversity can help maximize the recovery of overall biodiversity and ensure that unique species assemblages are re-established.
What is the concept of 'biotic homogenization'?
Biotic homogenization is the process by which the biological communities in different places become more similar over time, often due to human activities. It results in a decrease in beta diversity and a loss of unique local biotas.
How does 'habitat specialization' affect beta diversity?
Species that are highly specialized to particular habitats will contribute more to beta diversity, as they are likely to be found only in those specific conditions, leading to greater differentiation between communities.
What is the role of 'species pools' in determining beta diversity?
The regional species pool (the set of all species available to colonize local communities) sets the upper limit for beta diversity. The processes of dispersal and environmental filtering then determine which species from this pool establish in each local community.
How does 'sampling effort' affect beta diversity estimates?
Inadequate or unequal sampling effort can lead to biased estimates of beta diversity. If one site is undersampled, it might appear more dissimilar than it truly is, as many species might be missed.
What is the concept of 'beta-diversity decomposition'?
Beta-diversity decomposition is a method to break down total beta diversity into components related to species replacement (turnover) and species richness differences (nestedness), providing a more detailed understanding of the underlying ecological processes.
How does 'environmental stress' influence beta diversity?
Environmental stress can reduce the number of species that can tolerate certain conditions, leading to a decrease in alpha diversity. However, if different stressors affect different sites, it can also increase beta diversity by creating distinct, stress-adapted communities.
What is the significance of 'endemism' in beta diversity?
Endemic species (found only in a specific area) contribute significantly to beta diversity, as their unique presence makes a community distinct from others. Areas with high endemism are often conservation priorities.